jeudi 15 février 2018

Two 4-year PhD positions at the University of Groningen, the Netherlands

Project 1- Behavioural ecology & evolution: the role of phenotypic plasticity in species divergence
The Groningen Institute for Evolutionary Life Sciences (GELIFES) offers an NWO-funded PhD position for a research project on the role of phenotypic plasticity in speciation, using African cichlid fish as a model system.
Project description
Phenotypic plasticity is a key determinant of organismal performance, allowing rapid adjustment to environmental variation. In this project, we aim to investigate plasticity’s role in species divergence. On the one hand, plasticity may promote evolutionary differentiation by facilitating range expansion. On the other hand, plastic responses will weaken selection for genetic change, hampering adaptive evolution.
We study the role of visual plasticity in the evolution of cichlid fish species diversity. Variation in colour vision, among individuals, populations and species, is determined by both genetic factors and phenotypic plasticity. Since visually mediated behaviours influence both mating behaviour and ecological performance in cichlids, plasticity in colour vision could be important in divergent evolution. The PhD student will conduct experiments and analyses addressing the consequences of visual plasticity for both 1) macro-evolutionary processes and 2) individual behaviour and fitness. Approaches include behavioural experiments, molecular analysis of visual pigments (sequencing, qPCR, HPLC) and phylogenetic comparative analyses.
The project is a collaboration between Dr Martine Maan, Prof dr Rampal Etienne and Dr Louis van der Zande. The PhD student will join a lively and international team of researchers and (PhD) students working on (cichlid) behavioural ecology and evolution, supported by laboratory technicians and animal caretakers.
Qualifications
We are looking for an enthusiastic and talented PhD candidate with a master's degree (or equivalent) in Biology, preferably with a strong interest in behavioural ecology, evolution and biodiversity. The ideal candidate has experience with behavioural observation, animal handling and molecular genetics techniques. We also ask for a collaborative mindset, and excellent communication and writing skills (also in English). Other helpful competences include statistical proficiency (e.g. in the R software environment), possession of a driving licence, and a license to perform animal experiments (art. 9 Wet op de dierproeven (the Dutch Experiments on Animal Act)) or the willingness to obtain these.
For more information and application procedure, see:
Project 2 - Biology & philosophy: inheritance of acquired characters
The Young Academy of Groningen (YAG) was established in 2016 to bring together young researchers from diverse academic backgrounds. The YAG stimulates interdisciplinary research, contributes to internationalization and science policy, and promotes public outreach. We offer an interdisciplinary PhD scholarship for a project at the interface of biology and philosophy.

Project description
How do living organisms transmit their characteristics to the next generation? The textbook answer is: by passing on their DNA, which encodes all heritable traits. In recent years, however, biologists have discovered that this is not the whole story. Evidence is accumulating that environmental influences during the lifetime of an individual, such as nutrition or stress, can influence the phenotype of their progeny across several generations. This phenomenon reminds us of the concept of ‘inheritance of acquired characters’, already formulated in 1809 by the French naturalist Jean Baptiste de Lamarck in his Philosophie Zoologique. Lamarck’s ideas have often met with skepticism, but recent developments in molecular genetics have inspired a re-evaluation of his legacy. Indeed, we seem to witness a renaissance of Lamarckian thought.
In this project, we aim to explore the Lamarckian nature of recent discoveries in the transmission of inherited traits, and in this way to explore what modern biology can, or cannot, learn from the original ideas of Lamarck, whether or not the works of Lamarck offer lessons that change the way we think about evolution. This project will contribute to that debate by bringing together the disciplines of evolutionary biology, and the history of science and philosophy.
The project is a collaboration between Dr. Martine Maan (www.rug.nl/staff/m.e.maan) and Dr. Han Thomas Adriaenssen (www.rug.nl/staff/h.t.adriaenssen). 
Qualifications
The ideal candidate has a background in both biology and the history of science or philosophy. Alternatively, the candidate will have a background in one of these disciplines, and be prepared to be trained in the other as a part of their PhD trajectory. 
For more information and application procedure, see: